1. The Francis Crick Institute 1 Cellular signalling and cytoskeletal function laboratory , , 1 Midland Road, London, NW1 1AT , UK
2. London Research Institute, Cancer Research UK 2 , 44 Lincoln's Inn Fields, London, WC2A 3PX , UK
3. Imperial College 3 Department of Infectious Disease , , London W2 1PG , UK