1. North Carolina State University 1 Department of Biological Sciences , , Raleigh, NC 27695-7633 , USA
2. Center for Human Health and the Environment 2 , Department of Population Health and Pathobiology , , Raleigh, NC 27607 , USA
3. College of Veterinary Medicine, North Carolina State University 2 , Department of Population Health and Pathobiology , , Raleigh, NC 27607 , USA
4. University of North Carolina 3 Department of Neurology , , Chapel Hill, NC 27599 , USA