1. Department of Anatomy and Developmental Biology, Graduate School of Medical Science, Kyoto Prefectural University of Medicine, Kyoto, Japan
2. Department of Pathology and Cell Regulation, Graduate School of Medical Science, Kyoto Prefectural University of Medicine, Kyoto, Japan
3. Department of Anatomy and Cell Biology, Gunma University Graduate School of Medicine, Gunma, Japan
4. Developmental Genetics Group, Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan