1. University of North Carolina McAllister Heart Institute, UNC-Chapel Hill, Chapel Hill, NC 27599-3280, USA.
2. Department of Genetics, UNC-Chapel Hill, Chapel Hill, NC 27599-3280, USA.
3. Department of Molecular Biology. Princeton University, Princeton, NJ 08544, USA.
4. Department of Biology, UNC-Chapel Hill, Chapel Hill, NC 27599-3280, USA.
5. Section of Molecular Cell and Developmental Biology, University of Texas at Austin, Austin, TX 78712, USA.
6. Howard Hughes Medical Institute & Section of Molecular Cell and Developmental Biology, The University of Texas at Austin, Austin, TX 78712, USA.