1. Wellcome Trust/Cancer Research UK Gurdon Institute and Department of Genetics,Tennis Court Road, Cambridge CB2 1QR, UK
2. Department of Experimental Embryology, Polish Academy of Science, Jastrzebiec 05-552, Poland
3. Développement des Vertébrés, Institut Jacques Monod,UMR7592 CNRS Universités Paris 6, 7. 2, place Jussieu, 75005 Paris,France
4. Department of Microbiology, Shiga University of Medical Science, Otsu, Shiga 520-2192, Japan