1. University of Pennsylvania Perelman School of Medicine, Department of Cell and Developmental Biology, Philadelphia, PA 19104, USA
2. Biozentrum, University of Basel, Spitalstrasse 41, 4056 Basel, Switzerland
3. The Francis Crick Institute, 1 Midland Road, London NW1 1AT, UK
4. Center of Regenerative Medicine, Washington University School of Medicine, St. Louis, MO 63110, USA
5. Department of Cell and Developmental Biology, University College London,London WC1E 6BT, UK