1. Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, PA 19104-6058, USA.
2. Division of Basic Science, Fred Hutchinson Cancer Research Center Seattle, WA 98109, USA.
3. Molecular Neurobiology Program, The Helen L. and Martin S. Kimmel Center for Biology and Medicine at the Skirball Institute of Biomolecular Medicine, New York University Medical School, 540 First Avenue, New York, NY 10016, USA.