1. Department of Biology, University of Maryland, College Park, MD 20742, USA
2. Centre of Ecology, Evolution and Biogeochemistry , EAWAG Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland
3. Institute of Ecology and Evolution , University of Bern, 3012 Bern, Switzerland
4. Queensland Brain Institute, University of Queensland, Brisbane 4072 QLD, Australia