1. RIKEN Center for Integrative Medical Sciences 1 Laboratory of Functional Non-coding Genomics , , 1-7-22 Suehiro-cho, Tsurumi-ku, Yokohama, Kanagawa 230-0045 , Japan
2. Precursory Research for Embryonic Science and Technology (PRESTO) 2 Japan Science and Technology Agency (JST) , , Saitama 332-0012 , Japan
3. RIKEN Center for Integrative Medical Sciences 3 Laboratory of Developmental Genetics , , 1-7-22 Suehiro-cho, Tsurumi-ku, Yokohama, Kanagawa 230-0045 , Japan
4. Graduate School of Medicine, Chiba University 4 Department of Cellular and Molecular Medicine , , 1-8-1 Inohana, Chuo-ku, Chiba 260-8670 , Japan