1. University of Michigan Medical School 1 Department of Cell and Developmental Biology , , Ann Arbor, MI 48109 , USA
2. University of Michigan College of Engineering 2 Department of Biomedical Engineering , , Ann Arbor, MI 48109 , USA
3. University of Michigan Medical School 3 Department of Internal Medicine, Gastroenterology , , Ann Arbor, MI 48109 , USA