1. Department of Developmental Genetics, National Institute of Genetics, Yata 1111, Mishima 411-8540, Japan.
2. Graduate University for Advanced Studies, Yata 1111, Mishima 411-8540, Japan.
3. Laboratory for Molecular Developmental Biology, Department of Biophysics, Kyoto University, Kitashirakawa-Oiwake-tyo, Sakyo-ku, Kyoto 606-8502, Japan.
4. Temasek Life Sciences Laboratory and Department of Biological Sciences, 1 Research Link, National University of Singapore, Singapore 117604, Singapore.