1. Laboratory of Neuroscience, Graduate School of Frontier Biosciences, Osaka University, Suita, Osaka 565-0871, Japan.
2. SORST, Japan Science and Technology, Kawaguchi, Saitama 332-0012, Japan.
3. Division of Behavior and Neurobiology, National Institute for Basic Biology,Myodaiji-cho, Okazaki 444-8585, Japan.
4. Department of Development and Differentiation, Institute for Frontier Medical Sciences, Kyoto University, Kyoto 606-8507, Japan.