1. Department of Biology, Faculty of Science, and Graduate School of Science and Technology, Chiba University, Yayoicho, Inageku, Chiba, Chiba 263-8522,Japan
2. Division of Biochemistry, Institute of Medical Science, University of Tokyo,Shirokanedai, Minatoku, Tokyo 108-8639, Japan
3. Division of Cancer Genomics, Institute of Medical Science, University of Tokyo, Shirokanedai, Minatoku, Tokyo 108-8639, Japan
4. PRESTO, Japan Science and Technology Corporation (JST), Japan
5. CREST, Japan Science and Technology Corporation (JST), Japan