1. Roger Williams University 1 Marine Biology and Environmental Science , , Bristol, RI 02809 , USA
2. Providence College 2 Biology Department , , Providence, RI 02918 , USA
3. Whitman Center 3 , Marine Biological Laboratory, Woods Hole, MA 02543 , USA
4. Research and Development, Monterey Bay Aquarium Research Institute 4 , Moss Landing, CA 95039 , USA