1. Institute of Marine Science, University of Auckland 1 , Leigh Marine Research Laboratory, 160 Goat Island Road, Leigh, Auckland 0985 , New Zealand
2. The University of Florida 2 , Whitney Laboratory for Marine Bioscience, 9505 N Ocean Shore Blvd, St. Augustine, FL 32080 , USA
3. University of Auckland 3 Department of Statistics, Faculty of Science , , 38 Princes Street, Auckland 1010 , New Zealand