1. Research Institute for the Environment and Livelihoods, Charles Darwin University, Northern Territory 0909, Australia
2. Department of Zoology, University of Melbourne, Parkville, Victoria 3010, Australia
3. School of Biological Sciences, Monash University, Clayton Campus, Victoria 3800, Australia
4. School of Biological Sciences A08, University of Sydney, New South Wales 2006, Australia
5. School of Marine and Tropical Biology, James Cook University, Townsville, Queensland 4814, Australia