1. Developmental Biology Unit UMR 7009, UMPC Univ. Paris 06 and Centre National de la Recherche (CNRS), Observatoire Océanologique, 06230 Villefranche-sur-Mer, France
2. Institute of Cell and Molecular Bioscences, The Medical School, Framlington Place, University of Newcastle upon Tyne, Newcastle upon Tyne NE2 4HH, UK