1. MRC Centre for Reproductive Health/MRC Human Reproductive Sciences Unit, Queens Medical Research Institute, University of Edinburgh, 47 Little France Crescent, Edinburgh EH16 4TJ, UK
2. MRC Human Genetics Unit, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Crewe Road, Edinburgh EH4 2XU, UK
3. MRC-University of Glasgow Centre for Virus Research, Institute of Infection, Immunity, and Inflammation, College of Medicine, Veterinary Medicine and Life Sciences, Garscube Campus, University of Glasgow, Glasgow G61 1QH, UK