1. Department of Molecular and Cellular Biology, Harvard University, Cambridge,MA 02138, USA
2. Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, CA 94305, USA
3. Department of Microbiology, Columbia University College of Physicians and Surgeons, New York, NY 10032, USA
4. Wellcome Trust Center for Human Genetics, University of Oxford, Oxford OX3 7BN, UK