1. Murdoch Children's Research Institute, The Royal Children's Hospital, Flemington Road, Parkville, Victoria 3052, Australia
2. Department of Paediatrics, Faculty of Medicine, Dentistry and Health Sciences, University of Melbourne, Parkville, Victoria 3052, Australia
3. School of BioSciences, University of Melbourne, Parkville, Victoria 3052, Australia
4. Department of Anatomy and Developmental Biology, Monash University, Clayton, Victoria 3800, Australia