1. Department of Biological Sciences, California State University, East Bay,Hayward, CA 94542, USA
2. Department of Herpetology, California Academy of Sciences, San Francisco,CA 94103, USA
3. Department of Biology, Portland State University, Portland, OR 97207,USA
4. Zoology, School of Animal Biology M092, University of Western Australia,Crawley, Western Australia 6009, Australia