1. Howard Hughes Medical Institute and Abramson Family Cancer Research Institute,University of Pennsylvania School of Medicine, Philadelphia, PA 19104,USA
2. Abramson Family Cancer Research Institute, University of Pennsylvania School of Medicine, Philadelphia, PA 19104, USA
3. Department of Immunology, Medical Faculty/University Clinics, 89070 Ulm,Germany
4. Carl C, Icahn Center for Gene Therapy and Molecular Medicine, Mount Sinai School of Medicine, New York, NY 10029, USA