1. Department of Biology, McMaster University, 1280 Main St West, Hamilton,Ontario, Canada L8S 4K1
2. Rosenstiel School of Marine and Atmospheric Sciences, University of Miami,Miami, Florida 33149, USA
3. Bamfield Marine Sciences Centre, 100 Pachena Drive, Bamfield, British Columbia, Canada V0R 1B0
4. Department of Biology, University of Victoria, Victoria, British Columbia,Canada V8W 2N5