1. Division of Cell Biology and Biophysics, School of Biological Sciences, University of Missouri, Kansas City, MO 64110, USA
2. Hubrecht Institute-KNAW & University Medical Center Utrecht, 3584 CT Utrecht, The Netherlands
3. The Department of Cell Biology, UMC Utrecht, 3584 CX Utrecht, The Netherlands
4. Department of Biochemistry and Molecular Biophysics, Kansas State University, Manhattan, KS 66506, USA