1. Frontier Science Organization, Kanazawa University, 13-1 Takaramachi Kanazawa-shi, Ishikawa 920-8641, Japan
2. Graduate Program in Biophysics and Biochemistry, Graduate School of Science, University of Tokyo, 7-3-1 Hongo, Bunkyo-Ku, Tokyo 113-0033, Japan
3. Institute of Molecular and Cellular Biosciences, University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-0032, Japan
4. University of Massachusetts Medical School, Worcester, MA 01605, USA
5. PRESTO, JST, 4-1-8 Honcho Kawaguchi, Saitama 332-0012, Japan