1. 1Geological Survey of Canada, 3303 33rd Street NW, Calgary, Alberta T2L 2A7, Canada
2. 2Department of Earth, Energy, and Environment, University of Calgary, 2500 University Drive NW, Calgary, Alberta T2N 1N4, Canada
3. 3Department of Biology, Western Washington University, 516 High Street, Bellingham, Washington 98225, USA