1. 1Department of Earth and Environmental Sciences, University of Michigan, 1100 North University Avenue, Ann Arbor, Michigan 48109, USA
2. 2Department of Geology, University of Georgia, 210 Field Street, Athens, Georgia 30602, USA
3. 3Department of Ecology and Evolutionary Biology, University of Michigan, 1105 North University Avenue, Ann Arbor, Michigan 48109, USA