1. 1Papua New Guinea Forest Research Institute, Morobe Province, Papua New Guinea
2. 2Binatang Research Centre, Madang Province, Papua New Guinea
3. 3Faculty of Science, Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei
4. 4Department of Environmental Biology, Sapienza University of Rome, Rome, Italy
5. 5Papua New Guinea Food and Agricultural Organization, National Capital District, Papua New Guinea