1. MATTHEW L. HOLDING (matthewholding28@gmail.com) and ROBERT D. DENTON (denton.66@osu.edu) are joint first authors for this article and both are in the Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, Columbus, OH 43210.
2. AMY E. KULESZA (kulesza.5@osu.edu) and JUDITH S. RIDGWAY (ridgway.14@osu.edu) are at the Center for Life Sciences Education, The Ohio State University, Columbus, OH 43210.