1. Max Planck Institute for Evolutionary Biology, Department of Theoretical Biology, 24306 Plön, Germany
2. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544
3. Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, NJ 08544
4. Miller Institute for Basic Research in Science, University of California Berkeley, Berkeley, CA 94720
5. Department of Integrative Biology, University of California Berkeley, Berkeley, CA 94720