1. Department of Biology, Boston University, Boston, MA 02215
2. Graduate School of Agriculture, Kyoto University, Kyoto 606-8502, Japan
3. Research Faculty of Agriculture, Hokkaido University, Hokkaido 060-0809, Japan
4. Bioinformatics Graduate Program, Boston University, Boston, MA 02215
5. Department of Earth and Environment, Boston University, Boston, MA 02215