1. Research Center for Integrative Evolutionary Science, SOKENDAI (The Graduate University for Advanced Studies), Hayama 240-0193, Japan
2. Universal Biology Institute, The University of Tokyo, Bunkyo-ku 113-0033, Japan
3. CyberAgent, Inc., Shibuya-ku 150-0042, Japan
4. Department of Evolutionary Studies of Biosystems, SOKENDAI, Hayama 240-0193, Japan