1. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo 113-0033, Japan
2. Department of Plant Gene and Totipotency, Graduate School of Biostudies, Kyoto University, Kyoto 606-8502, Japan
3. Department of Bioscience and Biotechnology, Faculty of Environmental Sciences, Kyoto University of Advanced Science, Kyoto 621-8555, Japan