1. Department of Biological Sciences, Faculty of Science and Engineering, Chuo University, Tokyo 112-8551, Japan
2. Division of Developmental Biology, National Institute of Child Health and Human Development, NIH, Bethesda, MD 20892
3. Molecular and Cell Biology Department, Genetics, Genomics and Development Division, University of California, Berkeley, CA 94720
4. Department of Anatomy and Cell Biology, School of Medicine and Health Sciences, The George Washington University, Washington, DC 20037
5. Department of Molecular Pathobiology, New York University College of Dentistry, New York, NY 10010