1. European Molecular Biology Laboratory Australia Node in Single Molecule Science, School of Medical Sciences, University of New South Wales, Sydney 2052, NSW, Australia
2. Australian Research Council Centre of Excellence in Advanced Molecular Imaging, University of New South Wales, Sydney 2052, NSW, Australia
3. The Kennedy Institute of Rheumatology, University of Oxford, OX3 7FY Oxford, United Kingdom
4. Department of Mathematics and Statistics, Boston University, Boston, MA 02215
5. Center for Complex Biological Systems, University of California, Irvine, CA 92697
6. Sir William Dunn School of Pathology, University of Oxford, OX1 3RE Oxford, United Kingdom