1. Gembloux Agro-Bio Tech, University of Liege, 5030 Gembloux, Belgium
2. School of GeoSciences, University of Edinburgh, Edinburgh EH8 9YL, United Kingdom
3. Institute of Biological and Environmental Sciences, University of Aberdeen, Aberdeen AB24 3FX, United Kingdom
4. Departamento de Botânica, Universidade Federal de Minas Gerais, Belo Horizonte, 31270-901, Brazil
5. Evolutionary Biology and Ecology, Faculté des Sciences, Université Libre de Bruxelles, 1050 Brussels, Belgium