1. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA 98195
2. Department of Biology, University of Oklahoma, Norman, OK 73019
3. Department of Ecology and Evolutionary Biology, University of California, Irvine, CA 92697
4. Department of Evolution and Ecology, University of California, Davis, CA 95616
5. Department of Biological Sciences, Clemson University, Clemson, SC 29634