1. The Key Laboratory of Plant Development and Environment Adaptation Biology, Ministry of Education, School of Life Science, Shandong University, Qingdao 266237, China
2. School of Life Sciences, Southwest University, Chongqing 400715, China
3. Science and Technology Service Platform, Qilu University of Technology (Shandong Academy of Sciences), Jinan 250103, China
4. Maize Institute of Shandong Academy of Agricultural Sciences, Jinan, Shandong 250100, China
5. State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian, Shandong 271018, China