1. Cancer Research United Kingdom Scotland Centre, Institute of Genetics and Cancer, University of Edinburgh, Edinburgh EH4 2XU, United Kingdom
2. Medical Research Council Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Edinburgh EH4 2XU, United Kingdom
3. Division of Regulatory Genomics and Cancer Evolution (B270), German Cancer Research Center, Heidelberg 69120, Germany
4. Cancer Research United Kingdom Cambridge Institute, University of Cambridge, Cambridge CB2 0RE, United Kingdom
5. Medical Research Council Toxicology Unit, University of Cambridge, Cambridge CB2 1QR, United Kingdom
6. Department of Histopathology, Cambridge University Hospitals National Health Service Foundation Trust, Cambridge CB2 0QQ, United Kingdom