1. Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, CT 06269
2. Section of Entomology, University of Colorado Museum of Natural History, Boulder, CO 80309
3. Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen O 2100, Denmark
4. Departamento de Ecologia, Universidade Federal de Goiás, CP 131, 74.001-970 Goiânia, Goiás, Brazil