1. Department of Biology, Toho University, Funabashi 274-8510, Japan
2. Division of Forest & Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kyoto 606-8502, Japan
3. Division of Earth, Life and Environment, Graduate School of Human and Environmental Studies, Kyoto University, Kyoto 606-8501, Japan
4. Department of Human Ecology, Graduate School of Environmental and Life Science, Okayama University, Okayama 700-8530, Japan
5. Department of Biological Sciences, Florida Gulf Coast University, Ft. Myers, FL 33965