1. School of Life Sciences, Biomedical Pioneering Innovation Center, Peking University, Beijing 100871, People’s Republic of China
2. Changping Laboratory, Beijing 102206, People’s Republic of China
3. Joint Graduate Program of Peking-Tsinghua-National Institute of Biological Sciences, School of Life Sciences, Tsinghua University, Beijing 100084, People’s Republic of China
4. Beijing Advanced Innovation Center for Genomics, Peking University, Beijing 100871, People’s Republic of China