1. College of Horticulture, Fujian Agriculture and Forestry University, Fuzhou 350002, China
2. College of Life Sciences, Fujian Agriculture and Forestry University, Fuzhou 350002, China
3. School of Life Sciences, Lanzhou University, Lanzhou 730000, China
4. Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai 200032, China
5. Department of Biology, University of Pennsylvania, Philadelphia, PA 19104