1. Biomedical Pioneering Innovation Center, School of Life Sciences, Peking University, Beijing 100871, China
2. Academy for Advanced Interdisciplinary Studies, and Peking University–Tsinghua University–National Institute of Biological Sciences Joint Graduate Program, Peking University, Beijing 100871, China
3. Department of Physics and Graduate Institute of Biophysics, National Central University, Jhong-Li, Taoyuan 32001, Republic of China
4. School of Physics, Engineering and Technology, University of York, York YO10 5DD, United Kingdom
5. Department of Biology, University of York, York YO10 5DD, United Kingdom
6. Beijing Advanced Innovation Center for Genomics, Peking University, Beijing 100871, China