1. Flathead Lake Biological Station and Division of Biological Sciences, University of Montana, Polson, MT 59860
2. Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, China
3. Department of Land Resources and Environmental Sciences, Montana State University, Bozeman, MT 59717
4. Department of Ecosystem and Conservation Sciences, University of Montana, Missoula, MT 59812
5. Department of Integrative Biology, Oklahoma State University, Stillwater, OK 74078
6. Department of Biological Sciences, Great Lakes Research Center, Michigan Technological University, Houghton, MI 49931
7. Ecology, Evolution, and Organismal Biology program, University of Montana, Missoula, MT 59812