1. Faculty of Biological Sciences, University of Leeds, Leeds, LS2 9JT, United Kingdom
2. Leeds Omics, University of Leeds, Leeds, LS2 9JT, United Kingdom
3. Wellcome Centre for Human Genetics, University of Oxford, Oxford, OX3 7BN, United Kingdom
4. National Heart and Lung Institute, Imperial College London, London, SW3 6LY, United Kingdom