1. School of Natural Resource Management, Nelson Mandela University, Private Bag X6531, George, Western Cape, 6530 South Africa
2. Centre for Functional Biodiversity, School of Life Sciences, University of KwaZulu-Natal, 238 Mazisi Kunene Rd, Glenwood, Durban, 4041 South Africa
3. African Centre for Coastal Palaeoscience, Nelson Mandela University, P.O. Box 77000, Port Elizabeth, Eastern Cape, 6031 South Africa