1. Department of Veterinary Tropical Diseases, Faculty of Veterinary Science, Private Bag X04, Onderstepoort, 0110 South Africa
2. School of Biology and Environmental Sciences, University of Mpumalanga, Private Bag X11283, Nelspruit, 1200 South Africa
3. Department of Biological Sciences, University of Cape Town, Private Bag X3, Rondebosch, 7701 South Africa
4. Endangered Wildlife Trust, Private Bag X11, Modderfontein, 1645 South Africa
5. South African National Parks, P.O.Box 7814, Sonpark, 2016 South Africa