1. Cranfield Soil and Agrifood Institute, School of Water, Energy and Environment, Cranfield Univ.; Cranfield Bedfordshire MK43 0AL UK
2. Rothamsted Research; Harpenden Herts AL5 2JQ UK
3. James Hutton Institute; Craigiebuckler Aberdeen, Scotland AB15 8QH UK
4. Dep. of Soil and Environment; Swedish Univ. of Agricultural Sciences; Uppsala SE-750 07 Sweden