1. Sea Mammal Research Unit, Gatty Marine Laboratory, University of St. Andrews, St. Andrews, Fife, United Kingdom
2. North Carolina State College of Veterinary Medicine, Raleigh, North Carolina, USA
3. Institute of Zoology, Zoological Society of London, London, United Kingdom
4. Centre for Environment, Fisheries and Aquaculture Science, Burnham Laboratory, Burnham-on-Crouch, Essex, United Kingdom